The authors gratefully acknowledge Esther Martinez Aguilera for technical assistance. Houda Skouri-Gargouri and Hèla Trigui-Lahiani are thanked for their help and collaboration. This research was supported by Junta de Andalucía (grant CVI-7319), the Spanish Ministerio de Ciencia e Innovacion (grants BIO2013-47870 and BFU2015-67107) and Agencia Española de Cooperacion Internacional al Desarrollo (AECIDPCI-Med, AP/040009/11). GB had a postdoctoral position from Consejeria de Economia, Innovacion, Ciencia y Empresa, Junta de Andalucía.
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