This study was conducted without allocated funding. Catherine Greenfield is a Daphne Jackson Trust Research Fellow funded by NERC and the University of Leicester. Simon Gill is supported by a Royal Society Apex Award. Sam Poppe was supported by a frs-FNRS postdoctoral fellowship while at UlB and currently by a ULAM scholarship (NAWA Poland) and NCN Poland grant 2020/37/K/ST10/02447 at SRC PAS. The au- thors thank Tim Davis and Steffi Burchardt for discussions that benefitted the formulation of this paper, and thank the three anonymous reviewers and Editor—Jamie Farquharson—for review commentary that helped to improve the clarity of this paper.
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